My research studies important environmental and physiological influences including aggression, agonism, mating, opponent size, sex, resource availability, and hunger state.  My research also focuses on understanding the structure of the chemical, visual, mechanical, and social signals that animals receive in the natural environment. Recently, I have worked on characterizing how chemical signals travel in a local stream across variable substrates. I have also been recording social behavior and the ecological impact of animals that inhabit river and lake environments.